The werewolves of the world live in tight-knit gangs, or "dens," for protection from outsiders-and each other. Every major metropolis has one-to belong to a den is to have a family for eternity. However, Grace Holtz, next in line to lead the Seattle den called the Nameless, has had enough of living under the crushing weight of her den's expectations. Having fled to a small, rural town in Southeast Oregon, her goal is to blend in and be as "normal" as possible for the rest of her unnaturally long life.

But Marcus Bowen, a wolf from the UK-based Feóndulf den-and the closest thing Grace has ever had to a lover-has other plans. Reappearing thirty years after their affair came to an abrupt and bloody end, Marcus needs Grace. He needs her to return to Seattle and arrange an audience with Mama, the current leader of the Nameless and Grace's estranged grandmother. The leader of the Feóndulf and his heir have both been brutally murdered, and Marcus suspects that Grace and Mama are next.

Teaming up to hunt for the killer in the Emerald City, the pair slowly begin to realize their romance might not be as dead as they thought. However, as it becomes clear that the person they're looking for holds secrets about both their dens, Marcus and Grace must grapple with competing loyalties, conflicting desires, and ultimately decide what matters more-their dens or each other.